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What made the climate of northern Africa change from tropical to grasslands, and then to desert?

tilt of the earth's axis
the Nile River
Ice Ages
overpopulation of the region

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It was mostly the Ice Ages that made the climate of northern Africa change from tropical to grasslands, and then to desert, since only these massive changes and fluctuations in temperature could cause such a drastic shift. 
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User James Love
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